Red Pine Scale

Matsucoccus matsumurae

Images of this species:

Common look-alikes:


Red pine scale, now known more properly as pine bast scale, is an invasive insect that kills red pine trees in the United States. It is important not to confuse this with Matsucoccus macrocicatrices, a native pine bast scale that occurs on white pine trees.

Adult females are brick-red, wingless, soft-bodied insects with well-developed legs and antennae. They are approximately 1/16th”-3/16th” long, pear-shaped and coarsely wrinkled. Adult males are smaller (1/32nd”-1/16th”), two-winged, midge-like insects. Females deposit small, yellow eggs in a white, wooly ovisac. First-instar larvae resemble females, but are much smaller. Intermediate stage larvae are elliptical and lack legs and antennae. They resemble smooth, waxy pods, and have sparse tufts of white wool. Sub-adult males spin white, capsule-shaped, loosely woven cocoons. These are often visible on affected branches.

signs and symptoms

  • Foliage color changes slowly from light green to yellow to red
  • Masses of cottony white filaments become visible on the branches when infestations are heavy
  • Weakened trees may also be attacked by bark beetles causing rapid tree mortality

See image slideshow above for signs and symptoms.



Pine bast scale was first detected in North America in 1946 in Easton, Connecticut. It is thought to have been introduced to the United States on exotic pine trees planted at the NY World's Fair in 1939. Pine bast scale is native to Japan. It is considered an invasive species in China, Korea, Europe, and the United States. 


Pine bast scale attacks several Asian pine tree species as well as the North American red pine (Pinus resinosa).

Life Cycle

Pine bast scale has two generations per year. Eggs are laid in May, with first-instar larvae appearing by June. The larvae settle down and feed, usually under a bark scale. They enter the intermediate stage by mid-July. Pre-adult males emerge in August and spin a loosely woven cocoon in which they transform into winged true adults. Adult females emerge from the intermediate stage, mate, and lay eggs from August to September. The crawlers that hatch from this overwintering generation settle down and become dormant. They enter the intermediate stage beginning in April and rapidly develop into adults.

Like many scales, the insect spreads in the crawler stage by wind and as a hitchhiker, so the spread is generally slow. 

ecological threat

Research is ongoing, so it is premature to say that pine bast scale is the sole cause of red pine mortality in Vermont. Several shoot blight fungi are present in red pine stands and may play a role. Pine bast scale is cold sensitive, which may explain why it has been difficult to detect.

Management Options

Red pine scale is very difficult to detect. Suspect this insect if significant decline and red pine mortality are occurring, starting with dead shoots scattered in the crowns. Report suspect areas to 802-879-5687.

If red pine decline is occurring in parts of the state where the insect has been confirmed, the cause may likely be red pine scale. To prevent potential spread, use precautions when harvesting including:

  • Cutting stands in winter when the insect is not capable of moving on its own
  • Chipping tops so twigs and branches dry out more quickly
  • Ensuring ensure equipment is free of plant material before leaving the site.

Vermont Distribution

Red pine scale has not been confirmed in Vermont. 


photo credit

Alessandra Rung, Scale Insects, USDA APHIS ITP,

Allison Kanoti, Maine Forest Service,

information credit

Maine Pest Alert

Maine Forest Service

NH Bugs

Vermont 2015 Forest Insect and Disease Conditions Report

Vermont 2016 Forest Insect and Disease Conditions Report